Ghost Princess Perona is the former "Wild Zombies" and "Surprise Zombies Commander" of Thriller Bark before the collapse of Gekko Moriah's zombie army. She is able to produce ghosts from her body through the powers of the Horo Horo no Mi. She was one of the major antagonists of the Thriller Bark Arc, being one of the Mysterious Four, but has since begrudgingly cooperated with Roronoa Zoro.